Antispiral

The Anti-Spiral (アンチスパイラル, Anchi-Supairaru?) is the ultimate villain of Tengen Toppa Gurren-Lagann, and the central antagonistic presence for the final two story arcs. He first appears as a disembodied voice in Episode 24, but is formally introduced in Episode 26. His physical form is a chaos of arbitrary human concepts fluctuating on a black silhouette, with cartoonish gestures and exaggerated expressions.

Existence
Anti-Spiral is both the leader and collective mental manifestation of the "Anti-Spiral Race" after they seal themselves away to purge their race of Spiral energy. The highly advanced technology gives him extensive, almost supernatural powers, which manifest in the form of red energy. With these abilities he is able to manipulate physical and chemical laws (going as far as altering probability), which he does regularly both offensively and as a form of intimidation. The Anti-Spiral is the nemesis of Lordgenome, being responsible for Lordgenome's madness after explaining the nature of Spiral Nemesis to him.

Personality
Beyond what is known of the Anti-Spirals, little is known of the Anti-Spiral personally. He has the mannerisms of a scientist, suggesting he had a hand in both the creation of Anti-Spiral technology and the decision to undergo the millennia-long battle against the Milky Way's spiral beings. [3] The Anti-Spiral is very proud and holds deep contempt for spiral beings, specifically Simon, frustrated him that across all of time, spiral beings still resist limiting their populations and continue to defy the resolution of his race and their ideology.

History
See Anti-Spiral for more information

The Anti-Spiral sealed Team Dai-Gurren away in the Multi-Dimensional Labyrinth after they escaped the Spiral Sea. While Team Dai-Gurren was trapped, The Anti-Spiral was probing Nia for answers for her betrayal.

Gurren-Lagann appeared and rescued Nia. To counteract the newly created Tengen Toppa Gurren-Lagann, The Anti-Spiral created the Grand Zamboa. After the battle, The Anti-Spiral was defeated, destroying it, the Anti-Spiral Homeworld and the entire Anti-Spiral race.

Death
The Anti-Spiral is "killed" in Episode 27 by Simon and Nia in Lagann, killing the entire Anti-Spiral race in the process. But though he refuses to accept that his race were incorrect in restricting spiral energy, he dies asking Simon if Earth can take responsibility for protecting the universe. With his destruction, the "super spiral space" collapses and Team Dai-Gurren is returned to the physical world.

His death in the movie, Lagann-Hen, is different. Simon, instead so using Lagann Impact, fights The Anti-Spiral one on one, and is losing. Simon however, turns his hand into a Drill and stabs the Anti-Spiral with it, his death being the same after.