Guame

Guame the Immoveable (グァーム, Gwāmu?) is one of the Spiral King's Four Supreme Generals, who specializes in ground combat. His appearance is like that of a crocodile and armadillo and the oldest of the four generals, revealed in the parallel works to be originally an animal-like figure seen by Lordgenome's side as a parallel to Boota, living as long as or almost as long as the Spiral King has. In episode nine, he, along with the two other generals, reported the death of Thymilph to the Spiral King. He gives off the impression of one with a perverted mind and showed why when he made snide comments about the relationship between Thymilph and Adiane. Guame appears to merely respect the Spiral King, unlike the other Beastmen, who worship him as a god. When questioned by Cytomander after he boldly asked the king to let him go after Team Dai-Gurren, Guame explains his relation to King of being ages old (he later referred to Cytomander as a "pup of two-hundred-years age", implying that he is significantly older than Cytomander, enough for 200 years to seem like nothing). Succeeding in capturing the Dai-Gurren 's crew, Guame had Nia brought to him and reveals to her the truth he thought on why she was abandoned while attempting to offer to spare her life if she accepts to be leader of the "Beauty Village" where the Spiral King gets his consorts from. But Nia refused and Guame was forced to retreat when Team Dai-Gurren escaped and fight back.

A month after that, on the 6th day of Team Dai-Gurren's assault on Teppelin, Guame joins Cytomander in a two-pronged attack with both their battleships. After the Dai-Gunten is sunk, Guame uses Teppelin's power supply to have the Dai-Gundo create a massive tornado, in order to destroy Team Dai-Gurren and their reinforcements, but his plans are ruined when the Gurren-Lagann tunnels underneath the fortress to impale it with its drill, running Guame through in the process as the Dai-Gundo explodes.

Etymology
Guame's name is based on the DNA chemical guanine and the elemental Gnome. His Gunmen, Gember, is based on the mythical Black Tortoise, Genbu.

Beastman War Arc
Guame attacks Team Dai-Gurren by luring them into a trap with a fake distress signal from the beauty village. Simon regains his fighting spirit while freeing Team Dai-Gurren and defeats Guame. Guame and Cytomander combined forces to defend Teppelin from Team Dai-Gurren. Guame using Teppelin's power to make a giant tornado that destroying many new joines to Team Dai-Gurren, but Guame is defeated and killed by Gurren-Lagann when it drills underneath the Dai-Gundo and impales it.