Antispiral

The Anti-Spiral (アンチスパイラル, Anchisupairaru?) is the ultimate villain of Tengen Toppa Gurren-Lagann, and the central antagonistic presence for the final two story arcs. He first appears as a disembodied voice in Episode 24, but is formally introduced in Episode 26. His physical form is a chaos of arbitrary human concepts fluctuating on a black silhouette, with cartoonish gestures and exaggerated expressions.

Nature of Existence
Anti-Spiral is both the leader and the physical manifestation of the conciousness of the "Anti-Spiral race" after they sealed themselves away to purge their race of Spiral Energy. Their highly advanced technology gives him extensive, almost supernatural powers, which when used appear as red energy. With these abilities he is able to manipulate the laws of physics,and quantum mechanics (going as far as altering probability), which he wields regularly both offensively and as a form of intimidation. The Anti-Spiral is the nemesis of Lordgenome, being responsible for Lordgenome's madness after explaining the nature of the Spiral Nemesis to him.

Personality
Beyond what is known of the Anti-Spiral, little is known of the Anti-Spiral personally. He has the mannerisms of a scientist, suggesting he had a hand in both the creation of Anti-Spiral technology and the decision to undergo the millennia-long battle against the Milky Way's spiral beings. The Anti-Spiral is very proud and holds deep contempt for Spiral beings, specifically Simon, frustrated that across all of time, Spiral beings still resist the population limitations and continue to defy the resolution of his race and their ideology.

History
See Anti-Spiral for more information

The Anti-Spiral sealed Team Dai-Gurren away in the Extradimensional Labyrinth after they escaped the Death Spiral Field. While Team Dai-Gurren was trapped, The Anti-Spiral was probing Nia for answers for her betrayal.

Gurren Lagann appeared and rescued Nia as she was resisting the Anti-Spiral. To counteract the newly-created Tengen Toppa Gurren Lagann, the Anti-Spiral created the Granzeboma. After the battle, the Anti-Spiral was defeated, destroying it, the Anti-Spiral homeworld and the entire Anti-Spiral race.

Death
The Anti-Spiral is "killed" in Episode 27 by Simon and Nia in Lagann, killing the entire Anti-Spiral race in the process. But though he refuses to accept that his race were incorrect in restricting Spiral energy, he dies asking Simon if Earth can take responsibility for protecting the universe. With his destruction, the Super Spiral Universe collapses and Team Dai-Gurren is returned to the physical world.

His death in the movie, The Lights in the Sky are Stars, is different. Simon, instead of using Lagann Impact, fights the Anti-Spiral hand-to-hand, and is losing. Simon, however, turns the blood generated from the battle into a drill and pierces the Anti-Spiral with it, his death being the same afterwards.